Coverage-adjusted estimators for mark-recapture in heterogeneous populations.

Consideration of coverage yields a new class of estimators of population size for the standard mark-recapture model which permits heterogeneity of capture probabilities. Real data and simulation studies are used to assess these coverage-adjusted estimators. The simulations highlight the need for estimators that perform well for a wide range of values of the mean and coefficient of variation of the capture probabilities. When judged for this type of robustness, the simulations provide good grounds for preferring the new estimators to earlier ones for this model, except when the number of sampling occasions is large. A bootstrapping approach is used to estimate the standard errors of the new estimators, and to obtain confidence intervals for the population size.